It allowed the authors not only to identify three interconnected functional modules, but to also identify three new ciliopathy genes: Sang et al. 2011 identified ATXN10 and TCTN2, and Garcia-Gonzalo et al. 2011 identified TCTN1. The first module, the “nephronophthisis NPHP module” identified in Sang et al. 2011, was composed of NPHP1/nephrocystin-1, which was previously suggested to take part in the regulation of cargo and IFT ciliary entry[61]. Here, TCTN1 is linked to ciliopathy.