ATAD1 and infection: Intranuclear developing P. berghei wildtype and Δp52&p36 parasites were negative for UIS-4 peripheral staining, a marker for the presence of a PVM (Fig. 1b) and did not express MSP-1 at 52 hours post infection, as depicted by an intranuclear Δp52&p36 parasite (Fig. 1c).