For example, the glycoproteins of tospoviruses and rhabdoviruses and the minor capisd proteins of phytoreoviruses serve as viral ligands to mediate attachment of virions to receptors on the epithelial cells of the alimentary canal of the vector insect, a necessary step for virions to overcome midgut infection barriers of vector insect [2], [11]–[13], [38]–[40]. This evidence concerns the gene NR4A3 and infection.