However, in HIV-1 negative women maternal anemia was associated with KIR gene content polymorphisms in which women with subtype B1 (KIR2DS2−, KIR2DL2−, and KIR2DL3+) were less likely to have anemia and the association remained statistically significant after correction for multiple comparison (data not shown). Here, KIR3DL1 is linked to anemia (phenotype).