FGF4 and Severe short stature: For example, for proportionalheight at withers, we observe a large-effect QTL on CFA18 that we havepreviously shown corresponds to an fgf4 retrogene that accountsfor chrondrodysplasia or disproportional dwarfism in breeds such as corgis,basset hounds, and dachshunds [5], although we also find a novel QTL for heighton CFA20.