MUC5AC and colitis: We assessed the gene expression of several mucins that have been implicated in C. rodentium infection, and/or that are up-regulated in colitis, including the cell-surface mucins Muc1 and Muc3/17, and Muc13 [19], and the secreted non-gel forming mucin Muc4 that can be expressed by goblet cells [19], [55]; we also looked mucins that have gel-forming capacity, including the secreted gel-forming salivary and gastric mucins Muc19 [56] and Muc6 [57] respectively.