GLI1 and hydrops fetalis: In keeping with the more extensive ectopic HF morphogenesis in D2 than D4 transgenics, ectopic follicles in D2 mice that were heterozygous for the VDR contained cells expressing many transcriptional target genes of β-catenin/VDR complexes that are HF lineage markers, including K15, PADI3, S1003A, K31, K35, K71, K34, Gli1, and Dlx3 (Figure 4J, M, O, Q, S, U, W, Y, A').