We included HLA-DRB1*1501 in the analysis, even though we find no allelic association in our cohort, because previous studies have shown a greater risk for lupus in individuals who are compound heterozygotes for HLA-DRB1*0301- and HLA-DRB1*1501-containing haplotypes [25,28]. Here, HLA-DRB1 is linked to systemic lupus erythematosus.