MUC5AC and gallstones: Eleven regions were produced that exceeded the gFWER false positive cutoff (p-value = 0.000398; −log[p] = 3.400117), and seven of these regions fell within the range of traditionally identified QTL for gallstone formation or mucin accumulation, which is considered a precursor to gallstone formation (Table 1; D. Q. Wang et al. 1997).